Gavin Henson was ruled out on Friday of Wales' Six Nations opener against Scotland with a calf muscle injury.

Recalled to the team for the first time since Wales captured last season's grand slam, the centre was injured in training ahead of Sunday's match at Murrayfield.

Henson was sidelined for Wales' tour of South Africa and the November home internationals with an Achilles tendon injury.

His absence against the Scots means a recall for Tom Shanklin with his Cardiff Blues teammate Jamie Roberts moving from outside to inside centre.

Andrew Bishop will take Shanklin's place on the bench.

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